Adaptation of the Hashira Training Arc. The Hashira, the Demon Slayer Corps' highest ranking swordsmen and members. The Hashira Training has begun in order to face the forthcoming battle against Muzan Kibutsuji. Each with their own thoughts and hopes held in their hearts, a new story for Tanjiro and the Hashira begins.

Following the faceless god Hitogami's advice seems to have worked wonders for Rudeus Greyrat. After enrolling into the University of Magic as he was told, Rudeus reunites with his childhood friend Sylphiette, who put a valiant effort into curing his condition. The two grow ever closer together and decide to host a wedding party, inviting the friends they have made over the years to announce and formalize their relationship. For all his recent blessings, however, Rudeus' troubles are far from over. The research he is helping Shizuka Nanahoshi conduct hits a bottleneck, sending her into a deep slump much like he experienced in his previous life. Furthermore, a letter from his father, Paul, brings complications to Rudeus' relationships, and Sylphiette still knows next to nothing about his real background. In the face of these issues, Rudeus will have to apply the lessons he has learned in this new world to navigate through the challenges that come with living a life to its fullest.

With the ensemble contest behind them, the members of the Kitauji High School concert band now aim to win a gold medal at the national competition. For third-year club president Kumiko Oumae and her friends, it is their last chance before graduation. Filled with anticipation and anxiety, Kumiko worries if she can successfully lead the band. Amid her struggles, Kumiko encounters transfer student Mayu Kuroe. From her gentle demeanor to her talented euphonium skills, Mayu was a member of the Seira Girls' School concert band—a regular at nationals. While Mayu is friendly, Kumiko cannot help but have an indescribable feeling from their meeting. As new members arrive and plans are set in motion, the club, with an insatiable hunger for a gold medal, learns what it truly takes to propel themselves to reach the national level.

For the past 13 years, the Nefarious Monster Army have appeared beneath their floating fortress every Sunday to advance their goal of conquering Earth. Luckily, the Dragon Keepers are here to save the day! Having defeated the majority of the monsters during their initial invasion, the heroes now routinely show up to clear away any remnants of resistance. Unbeknownst to the public, however, every fight beyond the first has been a deliberate show put on by both sides: the Dragon Keepers reap the fame and prestige from fighting imaginary enemies, while the surviving weakest monsters, altogether called Dusters, are allowed to live for another day. Sentouin D, a Duster tired of living this life of shame, leaves the fortress in an attempt to overturn his fate. With the unexpected help of the mysterious ranger Yumeko Suzukiri, he discovers the key to defeating the Dragon Keepers lies in their Divine Artifacts—ultimate weapons crucial to the rangers' transformation and unique powers. Teaming up with Suzukiri to infiltrate the garrison of Red Keeper Sousei Akabane, Sentouin D must defy all expectations to save his species from the cruel hands of humans.